Resources

Periodicals

Ballinger, Royce E., Julio A. Lemos Espinal, and Geoffrey R. Smith. "Reproduction in Females of Three Species of Crevice-dwelling Lizards (Genus Xenosaurus) from Mexico." Studies on Neotropical Fauna and Environment 35 (2000): 179-183.

Cooper, William E., Jr., Julio A. Lemos-Espinal, and Geoffrey R. Smith. "Presence and Effect of Defensiveness or Context on Detectability of Prey Chemical Discrimination in the Lizard Xenosaurusplatyceps." Herpetologica 54 (1998): 409-413.

Herrel, Anthony, Ed De Grauw, and Julio A. Lemos-Espinal. "Head Shape and Bite Performance in Xenosaurid Lizards." Journal of Experimental Zoology 290 (2001): 101-107.

Lemos-Espinal, Julio A., Geoffrey R. Smith, and Royce E. Ballinger. "Diets of Three Species of Knob-Scaled Lizards (genus Xenosaurus) from México." Southwestern Naturalist 48 (2003).

-. "Ecology of Xenosaurus grandis agrenon, A Knob-Scaled

Lizard from Oaxaca, Mexico." Journal of Herpetology 37 (2003): In press.

-. "Natural History of the Mexican Knob-scaled Lizard,

Xenosaurus rectocollaris." Herpetological Natural History 4 (1996): 151-154.

-. "Natural History of Xenosaurus platyceps, A Crevice-

dwelling Lizard from Tamaulipas, Mexico." Herpetological Natural History 5 (1997): 181-186.

-. "Neonate-Female Associations in Xenosaurus newmanorum: A Case of Parental Care in a Lizard?" Herpetological Review 28 (1997): 22-23.

-. "Thermal Ecology of the Crevice-dwelling Lizard,

Xenosaurus newmanorum." Journal of Herpetology 32 (1998): 141-144.

Macey, J. Robert, James A. Schulte II, Allan Larson, Boris S. Tuniyev, Nikolai Orlov, and Theodore J. Papenfuss. "Molecular Phylogenetics, tRNA Evolution, and Historical Biogeography in Anguid Lizards and Related Taxonomic Families." Molecular Phylogenetics and Evolution 12 (1999): 250-272.

Nieto-Montes de Oca, A., Jonathan A. Campbell, and O. Flores-Villela. "A New Species of Xenosaurus (Squamata: Xenosauridae) from the Sierra Madre del Sur of Oaxaca, Mexico." Herpetologica 57 (2001): 32-47.

Pérez Ramos, E., L. Saldaña de la Riva, and Jonathan A. Campbell. "A New Allopatric Species of Xenosaurus (Squamata: Xenosauridae) from Guerrero, Mexico." Herpetologica 56 (2000): 500-506.

Smith, Geoffrey R., Julio A. Lemos-Espinal, and Royce E. Ballinger. "Sexual Dimorphism in Two Species of Knob-Scaled Lizards (Genus Xenosaurus) from Mexico." Herpetologica 53 (1997): 200-205.

Geoffrey R. Smith, PhD

Gila monsters and Mexican beaded lizards

(Helodermatidae)

Class Reptilia Order Squamata Suborder Scleroglossa Family Helodermatidae

Large, stout, venomous lizards with distinctive, beadlike scales (osteoderms) on the dorsal surfaces of head, limbs, body, and tail

1

12-18 in (30-45 cm) snout-to-vent length; 14-39 in (35-100 cm) total length; 1.0-4.4 lb (450g-2 kg)

Number of genera, species

1 genus; 2 species

Hot desert, tropical deciduous forest

Conservation status

Vulnerable: 2 species

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Distribution

Desert regions of the southwestern United States and northwestern Mexico; Pacific drainages along the western slope of Mexico and southern Guatemala; and two Atlantic drainages in Chiapas and eastern Guatemala

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