Physical characteristics

These are small to very large terrestrial turtles with a high-domed shell in all but one species; columnar hind limbs, ele-

A Greek tortoise (Testudo graeca) hatchling in Morocco. (Photo by Animals Animals ©M. Linley. Reproduced by permission.)

phantine in appearance; forelimbs somewhat flattened and armored with large scales; and short and unwebbed toes, each with two or fewer phalanges.

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